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chr12:45,890,331-52,362,786

Best TWAS P=4.38e-29 · Best GWAS P=4.2e-39 conditioned to 0.979

Associated models

#	Study	Tissue	Gene	h2	eQTL R2	model	# weights	model R2	model R2 P	eQTL GWAS Z	TWAS Z	TWAS P	Top SNP corr	PP3	PP4	joint
1	CommonMind	Brain Pre-frontal Cortex	C12orf54	0.08	0.01	bslmm	456	0.02	5.9e-04	-1.358	7.0	1.9e-12	-0.06	0.14	0.06	FALSE
2	CommonMind	Brain Pre-frontal Cortex	DIP2B	0.11	0.10	enet	16	0.13	2.6e-15	9.993	10.7	1.3e-26	0.04	0.12	0.88	TRUE
3	CommonMind	Brain Pre-frontal Cortex	TMEM106C	0.13	0.06	enet	21	0.11	3.2e-13	4.211	6.4	1.2e-10	-0.03	1.00	0.00	FALSE
4	GTEx	Adipose Subcutaneous	LETMD1	0.06	0.01	enet	27	0.00	1.7e-01	-4.843	-6.5	8.4e-11	0.03	0.11	0.42	FALSE
5	GTEx	Adipose Subcutaneous	ATF1	0.07	0.05	enet	14	0.04	3.1e-04	-7.522	-8.5	1.5e-17	-0.04	0.77	0.18	FALSE
6	GTEx	Adipose Subcutaneous	SMAGP	0.17	0.00	enet	22	0.02	1.3e-02	-4.758	5.3	1.2e-07	0.01	0.09	0.04	FALSE
7	GTEx	Adipose Visceral Omentum	CERS5	0.10	0.01	enet	11	0.01	1.3e-01	-6.764	5.6	2.1e-08	0.05	0.16	0.05	FALSE
8	GTEx	Adrenal Gland	DIP2B	0.16	0.05	lasso	9	0.07	2.1e-03	-6.307	-6.8	1.2e-11	-0.05	0.25	0.04	FALSE
9	GTEx	Artery Aorta	ATF1	0.26	0.29	lasso	2	0.28	1.5e-15	-7.547	-7.5	5.0e-14	-0.05	1.00	0.00	FALSE
10	GTEx	Artery Coronary	ATF1	0.20	0.09	lasso	4	0.05	1.2e-02	-7.522	-7.3	2.3e-13	-0.05	0.24	0.13	FALSE
11	GTEx	Artery Tibial	LIMA1	0.20	0.04	lasso	4	0.08	5.7e-07	-2.956	5.3	1.1e-07	0.04	0.38	0.34	FALSE
12	GTEx	Artery Tibial	ATF1	0.24	0.26	lasso	7	0.25	2.7e-19	-7.505	-7.5	5.3e-14	-0.05	1.00	0.00	FALSE
13	GTEx	Artery Tibial	RP4-605O3.4	0.04	0.01	lasso	4	0.00	3.2e-01	8.868	8.1	5.5e-16	0.03	0.24	0.04	FALSE
14	GTEx	Brain Caudate basal ganglia	LIMA1	0.18	0.12	lasso	5	0.07	3.9e-03	8.586	7.9	3.1e-15	0.04	0.12	0.29	FALSE
15	GTEx	Brain Cerebellar Hemisphere	PFKM	0.29	0.09	enet	14	0.06	1.5e-02	-8.320	9.0	2.2e-19	-0.02	0.02	0.94	TRUE
16	GTEx	Brain Cerebellar Hemisphere	RP4-605O3.4	0.18	0.12	enet	14	0.10	1.5e-03	8.949	7.9	2.6e-15	0.02	0.09	0.44	FALSE
17	GTEx	Brain Cerebellar Hemisphere	RP11-302B13.5	0.27	0.03	lasso	8	0.11	8.3e-04	3.974	-6.7	2.8e-11	-0.05	0.09	0.31	FALSE
18	GTEx	Brain Cerebellum	CSRNP2	0.22	0.25	lasso	6	0.20	1.3e-06	-4.893	-5.1	3.1e-07	0.03	0.84	0.05	FALSE
19	GTEx	Brain Cerebellum	PFKM	0.20	0.07	lasso	10	0.07	4.2e-03	-6.495	7.1	9.6e-13	-0.02	0.04	0.85	FALSE
20	GTEx	Brain Putamen basal ganglia	CERS5	0.29	0.12	enet	36	0.21	1.2e-05	8.424	8.2	2.3e-16	0.06	0.44	0.15	FALSE
21	GTEx	Breast Mammary Tissue	ATF1	0.07	0.07	lasso	5	0.04	2.9e-03	-6.519	-5.6	1.6e-08	-0.06	0.16	0.17	FALSE
22	GTEx	Breast Mammary Tissue	ASB8	0.17	0.02	enet	12	0.02	1.9e-02	7.235	6.2	7.2e-10	-0.03	0.11	0.04	TRUE
23	GTEx	Breast Mammary Tissue (Female)	ATF1	0.04	0.03	enet	7	0.00	2.7e-01	-6.519	-6.8	1.5e-11	-0.05	0.03	0.35	FALSE
24	GTEx	Breast Mammary Tissue (Female)	CCDC65	0.11	0.02	lasso	3	0.03	5.1e-02	-4.503	6.8	8.7e-12	-0.05	0.05	0.28	TRUE
25	GTEx	Cells Transformed fibroblasts	DIP2B	0.16	0.08	lasso	3	0.08	6.2e-07	-5.271	-6.4	1.7e-10	-0.06	0.94	0.01	FALSE
26	GTEx	Cells Transformed fibroblasts	SMARCD1	0.08	0.07	enet	5	0.07	9.3e-06	8.968	-8.8	2.1e-18	-0.03	0.96	0.03	FALSE
27	GTEx	Cells Transformed fibroblasts	POU6F1	0.18	0.12	lasso	4	0.15	1.7e-11	-4.480	-5.5	3.1e-08	0.00	1.00	0.00	FALSE
28	GTEx	Esophagus Gastroesophageal Junction	ATF1	0.15	0.15	lasso	4	0.14	7.8e-06	-7.108	-7.7	1.8e-14	-0.04	0.66	0.06	FALSE
29	GTEx	Esophagus Mucosa	TUBA1B	0.08	0.02	enet	24	0.04	1.1e-03	-2.684	6.1	1.2e-09	-0.07	0.19	0.11	TRUE
30	GTEx	Esophagus Mucosa	RP11-386G11.10	0.10	0.02	enet	23	0.05	2.0e-04	-2.684	5.8	7.4e-09	-0.08	0.46	0.08	FALSE
31	GTEx	Esophagus Muscularis	ATF1	0.15	0.17	lasso	2	0.16	7.5e-10	-7.547	-7.5	6.5e-14	-0.04	1.00	0.00	FALSE
32	GTEx	Esophagus Muscularis	CERS5	0.15	0.07	enet	18	0.09	2.7e-06	-6.716	6.5	7.0e-11	0.05	0.93	0.06	FALSE
33	GTEx	Heart Atrial Appendage	CCNT1	0.11	0.05	lasso	2	0.03	1.1e-02	-6.019	-5.9	2.7e-09	0.04	0.17	0.04	FALSE
34	GTEx	Heart Atrial Appendage	CERS5	0.14	0.03	lasso	6	0.02	4.2e-02	8.946	8.1	6.1e-16	0.04	0.51	0.12	FALSE
35	GTEx	Heart Left Ventricle	RP11-96H19.1	0.29	0.04	enet	33	0.13	1.7e-07	4.296	5.2	2.0e-07	-0.48	0.51	0.40	FALSE
36	GTEx	Liver	RP1-228P16.1	0.19	0.08	lasso	7	0.06	7.1e-03	7.235	-5.2	1.7e-07	0.05	0.23	0.04	FALSE
37	GTEx	Lung	CSRNP2	0.11	0.00	enet	19	0.09	3.1e-07	-4.624	-5.8	6.9e-09	0.04	0.72	0.01	FALSE
38	GTEx	Lung	CERS5	0.07	0.06	enet	12	0.07	2.8e-06	-3.738	6.1	1.2e-09	0.06	0.98	0.00	FALSE
39	GTEx	Lung	LARP4	0.07	0.05	enet	15	0.05	1.8e-04	-2.627	-5.5	3.1e-08	-0.06	0.44	0.50	FALSE
40	GTEx	Muscle Skeletal	LETMD1	0.10	0.07	enet	6	0.07	9.4e-08	-6.966	6.7	2.8e-11	0.01	1.00	0.00	FALSE
41	GTEx	Muscle Skeletal	DIP2B	0.03	0.04	lasso	6	0.04	1.1e-04	-7.505	-7.4	1.1e-13	-0.04	0.89	0.02	FALSE
42	GTEx	Muscle Skeletal	ATF1	0.09	0.06	lasso	5	0.07	5.0e-07	-7.547	-8.3	1.1e-16	-0.05	0.57	0.42	FALSE
43	GTEx	Muscle Skeletal	TFCP2	0.05	0.01	lasso	2	0.00	1.2e-01	-7.009	-7.0	2.8e-12	-0.02	0.57	0.01	FALSE
44	GTEx	Muscle Skeletal	KANSL2	0.15	0.10	lasso	3	0.09	1.8e-09	5.890	5.7	1.5e-08	-0.03	1.00	0.00	FALSE
45	GTEx	Muscle Skeletal	ZNF641	0.19	0.24	enet	7	0.20	4.0e-19	-5.659	-5.9	3.8e-09	0.04	1.00	0.00	FALSE
46	GTEx	Nerve Tibial	CERS5	0.31	0.28	enet	20	0.36	2.2e-26	8.946	8.5	1.7e-17	0.03	1.00	0.00	FALSE
47	GTEx	Nerve Tibial	RP4-605O3.4	0.11	0.00	lasso	3	0.00	1.7e-01	8.868	7.7	1.7e-14	0.02	0.14	0.15	FALSE
48	GTEx	Prostate	CERS5	0.19	0.08	lasso	4	0.02	1.2e-01	9.009	9.4	4.9e-21	0.02	0.08	0.12	FALSE
49	GTEx	Skin Not Sun Exposed Suprapubic	PFKM	0.15	0.06	enet	21	0.13	8.4e-08	1.816	6.8	1.4e-11	-0.04	0.76	0.14	TRUE
50	GTEx	Skin Not Sun Exposed Suprapubic	ZNF641	0.15	0.11	enet	30	0.10	3.1e-06	4.051	6.4	1.4e-10	-0.02	0.98	0.01	FALSE
51	GTEx	Skin Sun Exposed Lower leg	SMARCD1	0.07	0.02	lasso	2	0.03	1.4e-03	-5.708	-8.0	1.3e-15	0.01	0.16	0.50	TRUE
52	GTEx	Skin Sun Exposed Lower leg	ATF1	0.07	0.07	lasso	4	0.08	6.1e-07	-6.500	-6.8	1.0e-11	-0.06	0.98	0.01	FALSE
53	GTEx	Skin Sun Exposed Lower leg	CERS5	0.09	0.06	lasso	4	0.07	9.4e-07	-4.638	5.8	8.6e-09	0.05	1.00	0.00	FALSE
54	GTEx	Skin Sun Exposed Lower leg	ZNF641	0.21	0.03	enet	17	0.08	6.5e-07	-8.162	8.5	1.9e-17	-0.01	0.33	0.66	FALSE
55	GTEx	Skin Sun Exposed Lower leg	RP11-96H19.1	0.23	0.15	lasso	4	0.14	8.7e-12	4.296	6.0	1.8e-09	-0.46	1.00	0.00	FALSE
56	GTEx	Spleen	ZNF641	0.42	0.21	lasso	11	0.28	5.4e-08	-5.211	5.4	7.1e-08	-0.04	0.67	0.06	FALSE
57	GTEx	Stomach	DIP2B	0.09	0.09	lasso	5	0.07	4.1e-04	-5.271	-5.9	2.7e-09	-0.04	0.19	0.08	FALSE
58	GTEx	Testis	SMARCD1	0.22	0.10	enet	22	0.10	2.6e-05	8.502	-8.5	1.8e-17	-0.02	0.73	0.04	FALSE
59	GTEx	Testis	SLC11A2	0.17	0.13	enet	11	0.12	9.2e-06	-6.937	-8.7	4.6e-18	-0.04	0.67	0.25	TRUE
60	GTEx	Testis	CERS5	0.12	0.06	lasso	6	0.07	4.0e-04	8.949	9.4	6.9e-21	0.02	0.16	0.19	FALSE
61	GTEx	Testis	ZNF641	0.16	0.07	lasso	2	0.10	4.2e-05	-7.637	8.5	2.1e-17	-0.04	0.15	0.66	FALSE
62	GTEx	Thyroid	LIMA1	0.08	0.06	enet	18	0.06	1.7e-05	9.298	6.3	3.8e-10	0.02	0.22	0.36	FALSE
63	GTEx	Thyroid	CERS5	0.09	0.04	lasso	9	0.04	3.7e-04	-4.638	6.6	3.0e-11	0.04	0.80	0.15	FALSE
64	GTEx	Thyroid	ZNF641	0.16	0.06	enet	26	0.13	2.0e-10	-1.752	7.1	1.6e-12	-0.05	0.79	0.21	FALSE
65	GTEx	Thyroid	RP11-579D7.2	0.14	0.02	enet	31	0.02	7.3e-03	-6.495	5.4	6.0e-08	0.02	0.01	0.94	FALSE
66	GTEx	Whole Blood	DIP2B	0.12	0.08	lasso	4	0.08	5.7e-08	-5.271	-5.8	5.7e-09	-0.05	0.96	0.01	FALSE
67	GTEx	Whole Blood	ATF1	0.07	0.09	lasso	4	0.07	3.5e-07	8.814	8.8	9.0e-19	-0.03	0.89	0.11	FALSE
68	GTEx	Whole Blood	RP4-605O3.4	0.04	0.04	lasso	4	0.01	1.5e-02	9.009	9.1	6.1e-20	0.02	0.12	0.57	FALSE
69	METSIM	Adipose	DAZAP2	0.06	0.05	lasso	4	0.05	1.8e-07	-5.573	-5.8	5.9e-09	0.03	1.00	0.00	TRUE
70	METSIM	Adipose	METTL7A	0.06	0.00	blup	363	0.03	1.7e-05	-5.004	5.5	4.7e-08	0.00	0.32	0.02	FALSE
71	METSIM	Adipose	SLC38A2	0.02	0.02	blup	341	0.01	5.8e-03	-10.150	9.2	4.2e-20	-0.88	0.04	0.93	FALSE
72	NTR	Blood	ARF3	0.03	0.03	bslmm	339	0.03	8.4e-09	6.249	-5.2	1.7e-07	-0.09	0.00	1.00	FALSE
73	NTR	Blood	ATF1	0.06	0.07	enet	35	0.07	4.0e-21	10.350	9.5	2.0e-21	0.02	0.01	0.99	FALSE
74	NTR	Blood	SLC38A1	0.12	0.07	lasso	6	0.08	8.4e-24	5.822	-5.7	1.4e-08	0.02	1.00	0.00	TRUE
75	ROSMAP	Brain Pre-frontal Cortex	AMIGO2	0.06	0.00	blup	436	0.01	1.3e-02	-0.097	-5.8	7.9e-09	0.15	0.03	0.64	FALSE
76	YFS	Blood	ARF3	0.11	0.09	bslmm	335	0.10	4.8e-30	6.249	-6.7	1.8e-11	-0.08	0.00	1.00	TRUE
77	YFS	Blood	ATF1	0.05	0.07	lasso	9	0.07	2.0e-21	10.267	10.5	5.2e-26	0.03	0.01	0.99	FALSE
78	YFS	Blood	DIP2B	0.36	0.27	bslmm	385	0.28	2.8e-90	-5.271	6.3	2.7e-10	0.04	1.00	0.00	FALSE
79	YFS	Blood	LIMA1	0.04	0.06	enet	31	0.06	3.5e-19	-5.895	-7.6	4.2e-14	-0.05	1.00	0.00	FALSE
80	YFS	Blood	PFKM	0.03	0.00	blup	523	0.01	2.5e-05	-9.255	6.3	2.3e-10	-0.07	0.01	0.96	FALSE
81	YFS	Blood	SLC38A2	0.02	0.01	bslmm	338	0.01	2.2e-04	4.046	6.3	2.4e-10	-0.58	0.20	0.69	FALSE
82	The Cancer Genome Atlas	Bladder Urothelial Carcinoma	SLC38A4	0.07	0.02	lasso	5	0.04	1.3e-04	8.508	-9.8	1.2e-22	0.20	0.03	0.17	TRUE
83	The Cancer Genome Atlas	Breast Invasive Carcinoma	C12orf62	0.01	0.00	blup	47	0.00	4.6e-02	-3.569	-6.6	3.2e-11	-0.04	0.02	0.48	FALSE
84	The Cancer Genome Atlas	Breast Invasive Carcinoma	LETMD1	0.02	0.01	blup	44	0.01	4.4e-03	-3.285	5.3	9.5e-08	0.03	0.04	0.68	FALSE
85	The Cancer Genome Atlas	Breast Invasive Carcinoma	SLC11A2	0.03	0.02	blup	69	0.02	3.0e-04	-4.759	-5.6	2.8e-08	-0.07	0.83	0.07	FALSE
86	The Cancer Genome Atlas	Breast Invasive Carcinoma	SMARCD1	0.02	0.03	lasso	1	0.03	3.7e-06	8.590	-8.6	8.7e-18	-0.03	0.07	0.91	FALSE
87	The Cancer Genome Atlas	Colon Adenocarcinoma	ATF1	0.10	0.05	enet	18	0.06	3.5e-04	-7.504	-6.6	4.4e-11	-0.04	0.28	0.62	FALSE
88	The Cancer Genome Atlas	Colon Adenocarcinoma	DIP2B	0.13	0.14	lasso	2	0.12	1.3e-07	-5.515	-5.9	4.0e-09	-0.06	0.86	0.08	FALSE
89	The Cancer Genome Atlas	Head and Neck Squamous Cell Carcinoma	C12orf62	0.02	0.01	blup	47	0.01	1.7e-02	-3.738	-6.6	4.7e-11	-0.04	0.03	0.42	FALSE
90	The Cancer Genome Atlas	Head and Neck Squamous Cell Carcinoma	LETMD1	0.05	0.04	enet	3	0.04	8.2e-06	-6.758	6.0	1.9e-09	0.01	0.01	0.99	FALSE
91	The Cancer Genome Atlas	Head and Neck Squamous Cell Carcinoma	ZNF641	0.04	0.03	lasso	1	0.03	5.9e-04	-6.495	6.5	8.3e-11	-0.02	0.04	0.27	FALSE
92	The Cancer Genome Atlas	Kidney Renal Clear Cell Carcinoma	LETMD1	0.03	0.00	blup	44	0.02	3.8e-03	-3.879	6.2	4.4e-10	0.04	0.01	0.27	FALSE
93	The Cancer Genome Atlas	Kidney Renal Clear Cell Carcinoma	LIMA1	0.08	0.10	blup	59	0.10	1.8e-11	9.298	-9.6	6.3e-22	-0.02	0.00	1.00	FALSE
94	The Cancer Genome Atlas	Kidney Renal Papillary Cell Carcinoma	C12orf62	0.08	0.01	blup	47	0.06	3.2e-04	-3.569	-5.7	1.1e-08	-0.04	0.20	0.66	FALSE
95	The Cancer Genome Atlas	Kidney Renal Papillary Cell Carcinoma	SLC38A2	0.10	0.06	blup	30	0.05	8.3e-04	-11.576	9.4	5.0e-21	-0.52	0.10	0.07	FALSE
96	The Cancer Genome Atlas	Brain Lower Grade Glioma	ASB8	0.08	0.01	blup	69	0.04	3.9e-05	2.153	6.2	5.6e-10	-0.06	0.16	0.72	FALSE
97	The Cancer Genome Atlas	Brain Lower Grade Glioma	ATF1	0.03	0.04	lasso	1	0.04	6.3e-05	10.299	-10.3	7.1e-25	-0.02	0.01	0.99	FALSE
98	The Cancer Genome Atlas	Brain Lower Grade Glioma	CELA1	0.07	0.07	lasso	3	0.07	2.5e-08	5.481	5.3	1.3e-07	-0.03	0.01	0.99	FALSE
99	The Cancer Genome Atlas	Brain Lower Grade Glioma	FAM186A	0.03	0.03	lasso	6	0.02	2.2e-03	-6.450	5.8	5.2e-09	0.05	0.07	0.06	FALSE
100	The Cancer Genome Atlas	Brain Lower Grade Glioma	SLC11A2	0.11	0.11	lasso	3	0.11	2.3e-12	-4.869	5.2	1.6e-07	-0.03	0.99	0.01	FALSE
101	The Cancer Genome Atlas	Lung Adenocarcinoma	DIP2B	0.09	0.02	enet	9	0.05	1.3e-06	-5.515	-5.4	6.1e-08	-0.03	0.34	0.35	FALSE
102	The Cancer Genome Atlas	Lung Adenocarcinoma	ZNF641	0.03	0.03	lasso	7	0.03	3.1e-04	-7.194	6.8	9.1e-12	-0.04	0.12	0.08	TRUE
103	The Cancer Genome Atlas	Lung Squamous Cell Carcinoma	LETMD1	0.03	0.02	lasso	4	0.02	1.9e-03	-6.954	6.7	1.6e-11	0.03	0.01	0.86	FALSE
104	The Cancer Genome Atlas	Lung Squamous Cell Carcinoma	ZNF641	0.03	0.00	blup	50	0.01	2.6e-02	-5.659	6.3	3.8e-10	-0.05	0.03	0.12	FALSE
105	The Cancer Genome Atlas	Prostate Adenocarcinoma	LETMD1	0.03	0.03	blup	44	0.02	7.4e-03	-7.045	7.5	4.9e-14	0.01	0.01	0.91	FALSE
106	The Cancer Genome Atlas	Prostate Adenocarcinoma	TFCP2	0.03	0.02	blup	76	0.02	1.4e-03	-4.549	6.1	1.4e-09	-0.02	0.01	0.86	FALSE
107	The Cancer Genome Atlas	Soft Tissue Sarcoma	SLC38A2	0.06	0.01	blup	30	0.03	5.1e-03	-11.367	11.2	4.4e-29	-0.89	0.02	0.67	TRUE
108	The Cancer Genome Atlas	Testicular Germ Cell Tumors	DIP2B	0.11	0.02	blup	79	0.02	4.5e-02	-6.340	-6.5	9.3e-11	-0.05	0.06	0.25	FALSE
109	The Cancer Genome Atlas	Testicular Germ Cell Tumors	SLC11A2	0.17	0.09	lasso	11	0.08	1.0e-03	-6.852	-5.9	4.2e-09	0.00	0.04	0.62	FALSE
110	The Cancer Genome Atlas	Thyroid Carcinoma	C12orf41	0.09	0.06	lasso	3	0.07	4.3e-07	5.613	-5.6	2.5e-08	0.03	0.02	0.98	FALSE
111	The Cancer Genome Atlas	Thyroid Carcinoma	C12orf62	0.06	0.02	blup	47	0.05	1.8e-05	-2.339	-5.7	1.6e-08	-0.04	0.04	0.94	FALSE
112	The Cancer Genome Atlas	Thyroid Carcinoma	SLC38A1	0.04	0.03	blup	87	0.01	1.4e-02	-0.999	5.4	6.8e-08	-0.23	0.06	0.07	FALSE